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- Expression of the GBGT1 gene and the Forssman antigen in red blood cells in a Palestinian populationPublication . Abusibaa, Wafa Ali; Srour, Mahmoud A.; Moslemi, Ali-Reza; Svensson, Lola; Jesus, Carlos; Mendes, Fernando 1973-; Hesse, CamillaThe Forssman antigen (FORS1 Ag) is expressed on human red blood cells (RBCs). We investigated its presence on RBCs from Palestinian subjects and Swedish subjects by serological testing and by sequencing part of exon 7 of the GBGT1 gene, which encodes Forssman synthase.